
The Ghosts individually make their own way into Bolivia. The Ghost team consist of Ghost Leader "Nomad", Tactical Gunner "Midas", Engineer "Holt", Sniper "Weaver". The United States Army dispatches a fireteam of elite special operations unit called "Ghosts" to destroy the cartel and reveal the connection between Santa Blanca and the local government. As a result, the United States initiates Operation Kingslayer, a joint operation between the CIA, DEA and JSOC. embassy in La Paz and its intended target, DEA agent Ricardo "Ricky" Sandoval, is kidnapped, tortured and then killed by the Santa Blanca. The last straw comes when a bomb targets the U.S. The rise in power of this drug cartel concerns the United States Government, as Santa Blanca's influence has begun to spread beyond Bolivia. The country has become increasingly unstable as the Santa Blanca, a previously minor Mexican drug cartel, gains more power and influence within the county to the point where Bolivia has become a narco-state and the world's largest producer of cocaine. The game takes place in Bolivia in July 2019. According to the creative director of the game, the AI of the game will be unscripted and will have their "own motivations and agendas". Weapons and gear can be upgraded as well. The playable character can be customized, and loot found on enemies' corpses can be equipped by player characters.

Players can also gain experience points to level up. These interactions require strategy, as interactions will bring consequences and impact the game's world, and change how players can achieve their objectives. When not completing missions, players can interact, and build friendly or hostile relationships with other non-playable characters, such as the citizens, officials or the rebels of the world. Players can grab enemies at close range with one hand for defense, while using the other hand to shoot.
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Between missions, players are free to explore the game's world, and the game will feature outposts that can be taken down by players. Players are allowed to use multiple ways to complete objectives, such as utilizing stealth, melee combat, or using the long-ranged or short-ranged weapons provided in the game. Players can parachute from a helicopter, walk overland, or drive towards their objectives.

When completing missions, players can reach the location where the mission starts through a variety of ways. Wildlands will feature side-missions, unlike its predecessors. A variety of vehicles, such as dirt bikes, helicopters and dune buggies, will feature in the game. Players are tasked with making observations before carrying out missions. Completing missions during daytime allows players to spot enemies easily, while completing missions at night grants players a tactical advantage, as the night offers players better concealment. The game will be the first entry to feature an open world, which includes nine different types of terrain, such as: mountains, forests, desert, salt flats, and will introduce a dynamic weather system as well as a day-night cycle. The game will not feature the futuristic setting used in Advanced Warfighter and Future Soldier, but will instead adopt a modern-day setting, similar to the original Tom Clancy's Ghost Recon.

Players play as members of the Ghosts, a "fictional" elite special operations team in the United States Army. Tom Clancy's Ghost Recon Wildlands is an open world, tactical shooter utilising a third person perspective with an optional First-person view for gun aiming.
